Hi,
I said, it is the landing gear. Think of the actuators that move the gear. There is a big difference in their inside volume between extended (rod out) and retracted (rod in).
As I said, it is interesting why we see it only in #4 (actuates the body gear) and not also in #1 (actuates the nose and wing gears) and why there is more fluid in the reservoir when it should be less (when the gear is down). I think this question was asked before.
Hyd #1 and #4 also actuate the flap power packages but, in this case, it is only an hydraulic motors so there is a use of a lot of fluid (this is why #1 and #4 demand pumps run when flaps are out of up or 1, or when in transit – an airline option) but there is no real change to the volume.
Note that the quantity display shows a relative number, not a volume. There is more fluid in reservoirs 1 and 4 than 2 and 3.
Cheers,